In a list containing a series of items, separate the items with serial commas. However, a serial comma should not precede the conjunction before the final item (in other words, write a, b and c and not a, b, and c). But if a comma would make the meaning clearer, use it – especially where one of the items in the list is already joined by and.

Small_OK Specialist subjects include teaching, research and development, and business applications.

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